Pairing impressive reach with impressive speed, the Nikon AF-S NIKKOR 400mm f/2.8E FL ED VR is a super telephoto lens characterized by its bright f/2.8 maximum aperture and sophisticated optical, autofocus, and VR systems. Helping to achieve a high degree of clarity, sharpness, and color accuracy, the optical design uses two extra-low dispersion elements and two fluorite elements, which also contribute lessening the overall weight. Individual elements also feature both Nano Crystal and Super Integrated Coatings, which control lens flare and ghosting for improved contrast when working in bright and backlit conditions. And for protection, a front fluorine-coated meniscus glass element that, when paired with the weather-resistant magnesium-alloy barrel, supports working in inclement conditions.

A Silent Wave Motor AF system produces fast, near-silent focusing performance that suits both stills shooting and movie recording, which can instantly be manually overridden by simply turning the manual focus ring. Further contributing to photographing in fast-paced situations and difficult lighting, VR (Vibration Reduction) image stabilization compensates for the effects of camera shake by up to four shutter speed steps. Additionally, an electromagnetic aperture mechanism is integrated into the lens design to provide greater exposure control stability that is especially beneficial when working with faster continuous shooting rates.